Barrio Amoros, C.L. et al. 2021. Current status and natural history of the critically endangered Variable Harlequin Toad (Atelopus varius) in Costa Rica. Reptiles & Amphibians. 28, 3 (Nov. 2021), 374–388. DOI:https://doi.org/10.17161/randa.v28i3.15818.