Barrio Amoros, C. L., Chaves, G. ., & Puschendorf, R. (2021). Current status and natural history of the critically endangered Variable Harlequin Toad (Atelopus varius) in Costa Rica. Reptiles & Amphibians, 28(3), 374-388. https://doi.org/10.17161/randa.v28i3.15818 (Original work published 2021)