Barrio Amoros, C.L., Chaves, G. and Puschendorf, R. (2021) “Current status and natural history of the critically endangered Variable Harlequin Toad (Atelopus varius) in Costa Rica”, Reptiles & Amphibians, 28(3), pp. 374–388. doi:10.17161/randa.v28i3.15818.