C. L. Barrio Amoros, G. . Chaves, and R. Puschendorf, “Current status and natural history of the critically endangered Variable Harlequin Toad (Atelopus varius) in Costa Rica”, RandA, vol. 28, no. 3, pp. 374–388, Nov. 2021, doi: 10.17161/randa.v28i3.15818.