Habitat Suitability Analysis for Mountain Lions (Puma Concolor) Recolonization/ Reintroduction in Minnesota
DOI:
https://doi.org/10.2478/eje-2018-0013Keywords:
Suitability analysis, Mountain Lion, Model builder, food model, security model, habitat modelAbstract
The mountain lion range once extended throughout the state of Minnesota. The breeding population has been greatly reduced with time, new roads, and timber harvesting, which have broken large tracts of contiguous forest into isolated patches that are too small and no longer suitable for the breeding mountain lion population. The objective of this study is to use suitability analysis to determine the most suitable habitat to conserve mountain lion populations threatened by habitat fragmentation. To attain our objective, we created three sub models that contribute to the overarching goal of the suitability model. A habitat sub model was developed for finding the best habitat, a food sub model for access to the maximum amount of food needed, and a security sub model focusing on the distance from houses, roads, and urban development. Using the Weighted Sum tool, the three sub models were combined to produce a suitability surface based on the trade-off of the preferences of the goals represented by each sub model. Our suitability model shows large areas of high-quality mountain lion habitat in the northern and north-eastern sections of the state. These areas contain favourable locations for mountain lion habitat, such as forested land cover, low-density populations, steep slopes, short distances to streams, and area unimpeded by major roads. The southern and western parts of the state are characterized by lower slopes, more agricultural land, grassland, developed land, and higher population density, which results in lower quality habitat. The twin cities have the worst mountain lion habitat.
References
Alam, M. S., Khan, J. A., Kushwaha, S. P. S., Agrawal, R., Pathak, B. J.,
& Kumar, S. (2014). Assessment of suitable habitat of near
threatened striped hyena (Hyaena hyaena Linnaeus, 1758) using
Remote Sensing and Geographic Information System. Asian
Journal of Geoinformatics, 14(2). Retrieved from http://www.
geoinfo.ait.ac.th/ajg/index.php/journal/article/view/139
Alam, M. S., Khan, J. A., Kushwaha, S. P. S., Agrawal, R., Pathak, B. J.,
& Kumar, S. (2014). Assessment of suitable habitat of near
threatened striped hyena (Hyaena hyaena Linnaeus, 1758) using
Remote Sensing and Geographic Information System. Asian
Journal of Geoinformatics, 14(2). Retrieved from http://www.
geoinfo.ait.ac.th/ajg/index.php/journal/article/view/139
Alexander, S. M., Logan, T. B., & Paquet, P. C. (2006). Spatio-temporal
co-occurrence of cougars (Felis concolor), wolves (Canis lupus)
and their prey during winter: a comparison of two analytical
methods. Journal of Biogeography, 33(11), 2001–2012. https://
doi.org/10.1111/j.1365-2699.2006.01564.x
Alexander, S. M., Logan, T. B., & Paquet, P. C. (2006a). Spatio-temporal
co-occurrence of cougars (Felis concolor), wolves (Canis lupus)
and their prey during winter: a comparison of two analytical
methods. Journal of Biogeography, 33(11), 2001–2012. https://
doi.org/10.1111/j.1365-2699.2006.01564.x
Alexander, S. M., Logan, T. B., & Paquet, P. C. (2006b). Spatio-temporal
co-occurrence of cougars (Felis concolor), wolves (Canis lupus)
and their prey during winter: a comparison of two analytical
methods. Journal of Biogeography, 33(11), 2001–2012. https://
doi.org/10.1111/j.1365-2699.2006.01564.x
Alverson, W. S., Waller, D. M., & Solheim, S. L. (1988). Forests Too Deer:
Edge Effects in Northern Wisconsin. Conservation Biology, 2(4),
348–358. https://doi.org/10.1111/j.1523-1739.1988.tb00199.x
Anderson, A.E., (1983). A critical review of literature on Puma (Felis
concolor). Special Report 54. Colorado Division of Wildlife, Fort
Collins, CO, USA.
Apps, C. D., & McLellan, B. N. (2006). Factors influencing the dispersion
and fragmentation of endangered mountain caribou populations.
Biological Conservation, 130(1), 84–97. https://doi.
org/10.1016/j.biocon.2005.12.004
Apps, C. D., & McLellan, B. N. (2006). Factors influencing the dispersion
and fragmentation of endangered mountain caribou populations.
Biological Conservation, 130(1), 84–97. https://doi.
org/10.1016/j.biocon.2005.12.004
Arundel, T., D. Mattson, & Hart. J. (2007). Movements and habitat selection
by mountain lions in the Flagstaff uplands. Pages 17–30
in D. J. Mattson, editor. Mountain lions of the Flagstaff uplands,
2003–2006 progress report. U.S. Geological Survey Washington,
D.C., USA.
Baron, J. S., Theobald, D. M., & Fagre, D. B. (2000). Management of
Land Use Conflicts in the United States Rocky Mountains. Mountain
Research and Development, 20(1), 24–27. https://doi.
org/10.1659/0276-4741(2000)020[0024:MOLUCI]2.0.CO;2
Beier, P. (1993). Determining Minimum Habitat Areas and Habitat Corridors
for Cougars. Conservation Biology, 7(1), 94–108. https://
doi.org/10.1046/j.1523-1739.1993.07010094.x
Beier, P., & Noss, R. F. (1998). Do Habitat Corridors Provide Connectivity?
Conservation Biology, 12(6), 1241–1252. https://doi.
org/10.1111/j.1523-1739.1998.98036.x
Beier, P., Majka, D. R., & Newell, S. L. (2009). Uncertainty analysis of
least-cost modeling for designing wildlife linkages. Ecological
Applications, 19(8), 2067–2077. https://doi.org/10.1890/08-
1898.1
Beier, P., Majka, D. R., & Newell, S. L. (2009). Uncertainty analysis of
least-cost modeling for designing wildlife linkages. Ecological
Applications, 19(8), 2067–2077. https://doi.org/10.1890/08-
1898.1
Bolgiano, C., (1995). Mountain Lion: An Unnatural History of Pumas and
People. Stack-pole Books, Harrisburg, PA, USA.
Burdett, C. L., Crooks, K. R., Theobald, D. M., Wilson, K. R., Boydston,
E. E., Lyren, L. M., … Boyce, W. M. (2010). Interfacing models of
wildlife habitat and human development to predict the future
distribution of puma habitat. Ecosphere, 1(1), 1–21. https://doi.
org/10.1890/ES10-00005.1
Burdett, C. L., K. R. Crooks, D. M. Theobald, K. R. Wilson, E. E.Boydston,
L. M. Lyren, R. N.
Busch, R. H. (2004). The Cougar Almanac: A Complete Natural History of
the Mountain Lion (The Lyons Press: Guildord, CT).
Callan, R., Nibbelink, N. P., Rooney, T. P., Wiedenhoeft, J. E., & Wydeven,
A. P. (2013). Recolonizing wolves trigger a trophic cascade in
Wisconsin (USA). Journal of Ecology, 101(4), 837–845. https://
doi.org/10.1111/1365-2745.12095
Callan, R., Nibbelink, N. P., Rooney, T. P., Wiedenhoeft, J. E., & Wydeven,
A. P. (2013). Recolonizing wolves trigger a trophic cascade in
Wisconsin (USA). Journal of Ecology, 101(4), 837–845. https://
doi.org/10.1111/1365-2745.12095
Červinka, J., Šálek, M., Padyšáková, E., & Šmilauer, P. (2013). The effects
of local and landscape-scale habitat characteristics and prey
availability on corridor use by carnivores: A comparison of two
contrasting farmlands. Journal for Nature Conservation, 21(2),
105–113. https://doi.org/10.1016/j.jnc.2012.11.004
Clevenger, A. P., Wierzchowski, J., Chruszcz, B., & Gunson, K. (2002a).
GIS-Generated, Expert-Based Models for Identifying Wildlife
Habitat Linkages and Planning Mitigation Passages. Conservation
Biology, 16(2), 503–514. https://doi.org/10.1046/j.1523-
1739.2002.00328.x
Clevenger, A. P., Wierzchowski, J., Chruszcz, B., & Gunson, K. (2002b).
GIS-Generated, Expert-Based Models for Identifying Wildlife
Habitat Linkages and Planning Mitigation Passages. Conservation
Biology, 16(2), 503–514. https://doi.org/10.1046/j.1523-
1739.2002.00328.x
Crooks, K.R. and Soulé, M.E. (1999) Mesopredator Release and Avifaunal
Extinctions in a Fragmented System. Nature, 400, 563-566.
http://dx.doi.org/10.1038/23028
Crooks, K. R. (2002). Relative Sensitivities of Mammalian Carnivores to
Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Crooks, K. R. (2002a). Relative Sensitivities of Mammalian Carnivores
to Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Crooks, K. R. (2002b). Relative Sensitivities of Mammalian Carnivores
to Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Crooks, K. R. (2002c). Relative Sensitivities of Mammalian Carnivores
to Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Culver, M., Johnson, W. E., Pecon-Slattery, J., & O’Brien, S. J. (2000).
Genomic ancestry of the American puma (Puma concolor).
Journal of Heredity, 91(3), 186–197. https://doi.org/10.1093/
jhered/91.3.186
Cumming, G. S. (2000). Using habitat models to map diversity: pan-
African species richness of ticks (Acari: Ixodida). Journal of
Biogeography, 27(2), 425–440. https://doi.org/10.1046/j.1365-
2699.2000.00419.x
Dickson, B. G., & Beier, P. (2002). Home-Range and Habitat Selection by
Adult Cougars in Southern California. The Journal of Wildlife Management,
66(4), 1235–1245. https://doi.org/10.2307/3802956
Dickson, B. G., Jenness, J. S., Beier, P., & Gehrt. (2005). Influence of vegetation,
topography, and roads on cougar movement in southern california. Journal of Wildlife Management, 69(1), 264–276.
https://doi.org/10.2193/0022-541X(2005)069<0264:IOVTAR>2
.0.CO;2
Drake, N. (2011). Story one: Genes, sightings retrace path of cougar’s
journey: Biologists track cat from the Black Hills to Connecticut.
Science News, 180(5), 5–6. https://doi.org/10.1002/
scin.5591800503
Forman, R. T. T. (1987). The Ethics of Isolation, the Spread of Disturbance,
and Landscape Ecology. In Landscape Heterogeneity and
Disturbance (pp. 213–229). Springer, New York, NY. https://doi.
org/10.1007/978-1-4612-4742-5_12
Fortin, D., Beyer, H. L., Boyce, M. S., Smith, D. W., Duchesne, T., & Mao,
J. S. (2005). Wolves Influence Elk Movements: Behavior Shapes
a Trophic Cascade in Yellowstone National Park. Ecology, 86(5),
1320–1330. https://doi.org/10.1890/04-0953
Gehring, T. M., & Potter, B. A. (2005). Wolf habitat analysis in Michigan:
an example of the need for proactive land management for
carnivore species. Wildlife Society Bulletin, 33(4), 1237–1244.
https://doi.org/10.2193/0091-7648(2005)33[1237:WHAIMA]2.
0.CO;2
Gehring, T. M., & Potter, B. A. (2005). Wolf habitat analysis in Michigan:
an example of the need for proactive land management for
carnivore species. Wildlife Society Bulletin, 33(4), 1237–1244.
https://doi.org/10.2193/0091-7648(2005)33[1237:WHAIMA]2.
0.CO;2
Gilad, O., Wu, X. B., & Armstrong, F. (2013). Assessing the feasibility
for reintroducing desert bighorn sheep to Guadalupe Mountains
National Park: Habitat, migration corridors and challenges.
Applied Geography, 41, 96–104. https://doi.org/10.1016/j.apgeog.
2013.03.012
Gilad, O., Wu, X. B., & Armstrong, F. (2013a). Assessing the feasibility
for reintroducing desert bighorn sheep to Guadalupe Mountains
National Park: Habitat, migration corridors and challenges.
Applied Geography, 41, 96–104. https://doi.org/10.1016/j.apgeog.
2013.03.012
Gilad, O., Wu, X. B., & Armstrong, F. (2013b). Assessing the feasibility
for reintroducing desert bighorn sheep to Guadalupe Mountains
National Park: Habitat, migration corridors and challenges.
Applied Geography, 41, 96–104. https://doi.org/10.1016/j.apgeog.
2013.03.012
Girvetz, E. H., Thorne, J. H., Berry, A. M., & Jaeger, J. A. G. (2008). Integration
of landscape fragmentation analysis into regional planning:
A statewide multi-scale case study from California, USA.
Landscape and Urban Planning, 86(3), 205–218. https://doi.
org/10.1016/j.landurbplan.2008.02.007
Girvetz, E. H., Thorne, J. H., Berry, A. M., & Jaeger, J. A. G. (2008). Integration
of landscape fragmentation analysis into regional planning:
A statewide multi-scale case study from California, USA.
Landscape and Urban Planning, 86(3), 205–218. https://doi.
org/10.1016/j.landurbplan.2008.02.007
Glick, H. B. (2014). Modeling cougar habitat in the Northeastern
United States. Ecological Modelling, 285, 78–89. https://doi.
org/10.1016/j.ecolmodel.2014.02.005
Glick, H. B. (2014). Modeling cougar habitat in the Northeastern
United States. Ecological Modelling, 285, 78–89. https://doi.
org/10.1016/j.ecolmodel.2014.02.005
Gompper, M. E. (2002). Top Carnivores in the Suburbs? Ecological and
Conservation Issues Raised by Colonization of North-eastern
North America by CoyotesThe expansion of the coyote’s geographical
range may broadly influence community structure, and
rising coyote densities in the suburbs may alter how the general
public views wildlife. BioScience, 52(2), 185–190. https://doi.
org/10.1641/0006-3568(2002)052[0185:TCITSE]2.0.CO;2
Gray, M., Wilmers, C. C., Reed, S. E., & Merenlender, A. M. (2016a).
Landscape feature-based permeability models relate to puma
occurrence. Landscape and Urban Planning, 147, 50–58. https://
doi.org/10.1016/j.landurbplan.2015.11.009
Gray, M., Wilmers, C. C., Reed, S. E., & Merenlender, A. M. (2016b).
Landscape feature-based permeability models relate to puma
occurrence. Landscape and Urban Planning, 147, 50–58. https://
doi.org/10.1016/j.landurbplan.2015.11.009
Hansen, A. J., Rasker, R., Maxwell, B., Rotella, J. J., Johnson, J. D.,
Parmenter, A. W., … Kraska, M.P. V. (2002). Ecological Causes
and Consequences of Demographic Change in the New West.
BioScience, 52(2), 151–162. https://doi.org/10.1641/0006-
3568(2002)052[0151:ECACOD]2.0.CO;2
Harveson, L.A., Harveson, P.M. & Adams, R.W.(2003). Proceedings of
the Sixth Mountain Lion Workshop , Austin, Texas, USA.
Hénaux, V., Powell, L. A., Hobson, K. A., Nielsen, C. K., & LaRue, M. A.
(2011). Tracking large carnivore dispersal using isotopic clues
in claws: an application to cougars across the Great Plains.
Methods in Ecology and Evolution, 2(5), 489–499. https://doi.
org/10.1111/j.2041-210X.2011.00107.x
Hénaux, V., Powell, L. A., Hobson, K. A., Nielsen, C. K., & LaRue, M. A.
(2011). Tracking large carnivore dispersal using isotopic clues
in claws: an application to cougars across the Great Plains.
Methods in Ecology and Evolution, 2(5), 489–499. https://doi.
org/10.1111/j.2041-210X.2011.00107.x
Holmes, B. R., & Laundré, J. W. (2006). Use of open, edge and forest
areas by pumas Puma concolor in winter: are pumas foraging
optimally? Wildlife Biology, 12(2), 201–209. https://doi.
org/10.2981/0909-6396(2006)12[201:UOOEAF]2.0.CO;2
Hortal, J., Lobo, J. M., & Martín-piera, F. (2001). Forecasting insect
species richness scores in poorly surveyed territories: the
case of the Portuguese dung beetles (Col. Scarabaeinae).
Biodiversity & Conservation, 10(8), 1343–1367. https://doi.
org/10.1023/A:1016624500023
Hunter, R. D., Fisher, R. N., & Crooks, K. R. (2003). Landscape-level
connectivity in coastal southern California, USA, as assessed
through carnivore habitat suitability. Natural Areas Journal,
23(4), 302314.
Jenkins, M. (2003a). Prospects for Biodiversity. Science, 302(5648),
1175–1177. https://doi.org/10.1126/science.1088666
Jenkins, M. (2003b). Prospects for Biodiversity. Science, 302(5648),
1175–1177. https://doi.org/10.1126/science.1088666
Keeley, A. T. H., Beier, P., & Gagnon, J. W. (2016). Estimating landscape
resistance from habitat suitability: effects of data source and nonlinearities. Landscape Ecology, 31(9), 2151–2162. https://
doi.org/10.1007/s10980-016-0387-5
Kertson, B. N., Spencer, R. D., Marzluff, J. M., Hepinstall-Cymerman, J.,
& Grue, C. E. (2011). Cougar space use and movements in the
wildland–urban landscape of western Washington. Ecological
Applications, 21(8), 2866–2881. https://doi.org/10.1890/11-
0947.1
Knopff, A. A., Knopff, K. H., Boyce, M. S., & St. Clair, C. C. (2014). Flexible
habitat selection by cougars in response to anthropogenic development.
Biological Conservation, 178, 136–145. https://doi.
org/10.1016/j.biocon.2014.07.017
Knopff, A. A., Knopff, K. H., Boyce, M. S., & St. Clair, C. C. (2014). Flexible
habitat selection by cougars in response to anthropogenic development.
Biological Conservation, 178, 136–145. https://doi.
org/10.1016/j.biocon.2014.07.017
Kushwaha, S. P. S., Khan, A., Habib, B., Quadri, A., & Singh, A. (2004).
Evaluation of sambar and muntjak habitats using geostatistical
modelling. Current Science, 86(10), 1390–1400.
LaRue, M. A., & Nielsen, C. K. (2008). Modelling potential dispersal corridors
for cougars in midwestern North America using least-cost
path methods. Ecological Modelling, 212(3), 372–381. https://
doi.org/10.1016/j.ecolmodel.2007.10.036
LaRue, M. A., & Nielsen, C. K. (2008). Modelling potential dispersal corridors
for cougars in midwestern North America using least-cost
path methods. Ecological Modelling, 212(3), 372–381. https://
doi.org/10.1016/j.ecolmodel.2007.10.036
LaRue, M. A., & Nielsen, C. K. (2016). Population viability of recolonizing
cougars in midwestern North America. Ecological Modelling, 321,
121–129. https://doi.org/10.1016/j.ecolmodel.2015.09.026
LaRue, M. A., & Nielsen, C. K. (2016a). Population viability of recolonizing
cougars in midwestern North America. Ecological
Modelling, 321, 121–129. https://doi.org/10.1016/j.ecolmodel.
2015.09.026
LaRue, M. A., & Nielsen, C. K. (2016b). Population viability of recolonizing
cougars in midwestern North America. Ecological
Modelling, 321, 121–129. https://doi.org/10.1016/j.ecolmodel.
2015.09.026
LaRue, M. A., Nielsen, C. K., Dowling, M., Miller, K., Wilson, B., Shaw,
H., & Anderson, C. R. (2012). Cougars are recolonizing the midwest:
Analysis of cougar confirmations during 1990–2008. The
Journal of Wildlife Management, 76(7), 1364–1369. https://doi.
org/10.1002/jwmg.396
Laundré, J. W., & Loxterman, J. (2007). Impact of Edge Habitat on Summer
Home Range Size in Female Pumas. The American Midland
Naturalist, 157(1), 221–229. https://doi.org/10.1674/0003-
0031(2007)157[221:IOEHOS]2.0.CO;2
Lenton, S. M., Fa, J. E., & Val, J. P. D. (2000). A simple non-parametric GIS
model for predicting species distribution: endemic birds in Bioko
Island, West Africa. Biodiversity & Conservation, 9(7), 869–885.
https://doi.org/10.1023/A:1008980910283
Light, J. T. (1971). An ecological view of bighorn habitat on Mt. San Antonio.
Transactions of North American wild sheep Conference 1,
pp. 150-157.
Liu, S., Dong, Y., Cheng, F., Zhang, Y., Hou, X., Dong, S., & Coxixo, A.
(2017). Effects of road network on Asian elephant habitat and
connectivity between the nature reserves in Xishuangbanna,
Southwest China. Journal for Nature Conservation, 38, 11–20.
https://doi.org/10.1016/j.jnc.2017.05.001
Liu, S., Dong, Y., Cheng, F., Zhang, Y., Hou, X., Dong, S., & Coxixo, A.
(2017). Effects of road network on Asian elephant habitat and
connectivity between the nature reserves in Xishuangbanna,
Southwest China. Journal for Nature Conservation, 38, 11–20.
https://doi.org/10.1016/j.jnc.2017.05.001
Logan, K. A., & Irwin, L. L. (1985). Mountain Lion Habitats in the Big
Horn Mountains, Wyoming. Wildlife Society Bulletin (1973-
2006), 13(3), 257–262.
Logan, K. A., & Sweanor, L. L. (2001). Desert Puma: Evolutionary Ecology
And Conservation Of An Enduring Carnivore. Island Press.
Maehr, D. S., Land, E. D., Shindle, D. B., Bass, O. L., & Hoctor, T. S.
(2002). Florida panther dispersal and conservation. Biological
Conservation, 106(2), 187–197. https://doi.org/10.1016/S0006-
3207(01)00245-2
Marzluff, J. M., & Ewing, K. (2001). Restoration of Fragmented Landscapes
for the Conservation of Birds: A General Framework and
Specific Recommendations for Urbanizing Landscapes. Restoration
Ecology, 9(3), 280–292. https://doi.org/10.1046/j.1526-
100x.2001.009003280.x
Marzluff, J. M., & Ewing, K. (2008). Restoration of Fragmented Landscapes
for the Conservation of Birds: A General Framework and
Specific Recommendations for Urbanizing Landscapes. In Urban
Ecology (pp. 739–755). Springer, Boston, MA. https://doi.
org/10.1007/978-0-387-73412-5_48
Morrison, C. D., Boyce, M. S., Nielsen, S. E., & Bacon, M. M. (2014). Habitat
selection of a re-colonized cougar population in response to
seasonal fluctuations of human activity. The Journal of Wildlife
Management, 78(8), 1394–1403. https://doi.org/10.1002/
jwmg.799
Morrison, C. D., Boyce, M. S., Nielsen, S. E., & Bacon, M. M. (2014). Habitat
selection of a re-colonized cougar population in response to
seasonal fluctuations of human activity. The Journal of Wildlife
Management, 78(8), 1394–1403. https://doi.org/10.1002/
jwmg.799
Morrison, S. A., & Boyce, W. M. (2009). Conserving Connectivity: Some
Lessons from Mountain Lions in Southern California. Conservation
Biology, 23(2), 275–285. https://doi.org/10.1111/j.1523-
1739.2008.01079.x
Northrup, J. M., Pitt, J., Muhly, T. B., Stenhouse, G. B., Musiani, M., &
Boyce, M. S. (2012). Vehicle traffic shapes grizzly bear behaviour
on a multiple-use landscape. Journal of Applied Ecology, 49(5),
1159–1167. https://doi.org/10.1111/j.1365-2664.2012.02180.x
Noss, R. F., & Daly K. M. (2006). Incorporating connectivity into broad
scale conservation planning. Pages 587–619 in K. R. Crooks and
M. A. Sanjayan, editors. Connectivity conservation, Cambridge
University Press, Cambridge, United Kingdom
Noss, R. F., Quigley, H. B., Hornocker, M. G., Merrill, T., & Paquet, P.
C. (1996). Conservation Biology and Carnivore Conservation in
the Rocky Mountains. Conservation Biology, 10(4), 949–963.
https://doi.org/10.1046/j.1523-1739.1996.10040949.x
O′Neil, S. T., Rahn, K. C., & Bump, J. K. (2014). Habitat Capacity for Cougar
Recolonization in the Upper Great Lakes Region. PLOS ONE,
9(11), e112565. https://doi.org/10.1371/journal.pone.0112565
Pletscher, D. H., Ream, R. R., Boyd, D. K., Fairchild, M. W., & Kunkel, K. E.
(1997). Population Dynamics of a Recolonizing Wolf Population.
The Journal of Wildlife Management, 61(2), 459–465. https://
doi.org/10.2307/3802604
Pyare, S., Cain, S., Moody, D., Schwartz, C., & Berger, J. (2004). Carnivore
re-colonisation: reality, possibility and a non-equilibrium century
for grizzly bears in the southern Yellowstone ecosystem. Animal
Conservation Forum, 7(1), 71–77. https://doi.org/10.1017/
S1367943003001203
Restas, A. (2015). Drone Applications for Supporting Disaster Management.
World Journal of Engineering and Technology, 03(03),
316–321. https://doi.org/10.4236/wjet.2015.33C047
Ripple, W. J., & Beschta, R. L. (2004). Wolves and the Ecology of Fear: Can
Predation Risk Structure Ecosystems? BioScience, 54(8), 755–
766. https://doi.org/10.1641/0006-3568(2004)054[0755:WATE
OF]2.0.CO;2
Ripple, W. J., & Beschta, R. L. (2004a). Wolves and the Ecology of Fear:
Can Predation Risk Structure Ecosystems? BioScience, 54(8), 755–
766. https://doi.org/10.1641/0006-3568(2004)054[0755:WATE
OF]2.0.CO;2
Ripple, W. J., & Beschta, R. L. (2004b). Wolves and the Ecology of Fear:
Can Predation Risk Structure Ecosystems? BioScience, 54(8), 755–
766. https://doi.org/10.1641/0006-3568(2004)054[0755:WATE
OF]2.0.CO;2
Ripple, W. J., & Beschta, R. L. (2006a). Linking a cougar decline, trophic
cascade, and catastrophic regime shift in Zion National
Park. Biological Conservation, 133(4), 397–408. https://doi.
org/10.1016/j.biocon.2006.07.002
Ripple, W. J., & Beschta, R. L. (2006b). Linking a cougar decline, trophic
cascade, and catastrophic regime shift in Zion National
Park. Biological Conservation, 133(4), 397–408. https://doi.
org/10.1016/j.biocon.2006.07.002
Ripple, W. J., & Beschta, R. L. (2008). Trophic cascades involving cougar,
mule deer, and black oaks in Yosemite National Park. Biological
Conservation, 141(5), 1249–1256. https://doi.org/10.1016/j.
biocon.2008.02.028
Rosatte, R. (2011). Evidence Confirms the Presence of Cougars ( Puma
concolor ) in Ontario, Canada. The Canadian Field-Naturalist,
125(2), 116–125.
Saaty, T.L. (1980). The Analytical Hierarchy Process: Planning, Setting
Priorities, Resource Allocation. McGraw-Hill International Book
Co., New York.
Sanderson, E. W., Redford, K. H., Vedder, A., Coppolillo, P. B., & Ward,
S. E. (2002a). A conceptual model for conservation planning
based on landscape species requirements. Landscape and Urban
Planning, 58(1), 41–56. https://doi.org/10.1016/S0169-
2046(01)00231-6
Sanderson, E. W., Redford, K. H., Vedder, A., Coppolillo, P. B., & Ward,
S. E. (2002b). A conceptual model for conservation planning
based on landscape species requirements. Landscape and Urban
Planning, 58(1), 41–56. https://doi.org/10.1016/S0169-
2046(01)00231-6
Schamberger, M., & Krohn, W. (1982). Status of the Habitat Evaluation
Procedures. US Fish & Wildlife Publications. Retrieved from
https://digitalcommons.unl.edu/usfwspubs/48
Schipper, J., Chanson, J. S., Chiozza, F., Cox, N. A., Hoffmann, M., Katariya,
V., … Young, B. E. (2008). The Status of the World’s Land and
Marine Mammals: Diversity, Threat, and Knowledge. Science,
322(5899), 225–230. https://doi.org/10.1126/science.1165115
Shen, G., Feng, C., Xie, Z., Ouyang, Z., Li, J., & Pascal, M. (2008). Proposed
Conservation Landscape for Giant Pandas in the Minshan
Mountains, China. Conservation Biology, 22(5), 1144–1153.
https://doi.org/10.1111/j.1523-1739.2008.01038.x
Singer, F. J., Bleich, V. C., & Gudorf, M. A. (2000). Restoration of Bighorn
Sheep Metapopulations in and Near Western National Parks.
Restoration Ecology, 8(4S), 14–24. https://doi.org/10.1046/
j.1526-100x.2000.80062.x
Smith, T. S., & Flinders, J. T. (1992). Evaluation of mountain sheep habitat
in Zion National Park, Utah. Desert Bighorn Council Transactions,
36, 4-9.
Smith, T. S., Flinders, J. T., & Winn, D. S. (1991a). A HABITAT EVALUATION
PROCEDURE FOR ROCKY MOUNTAIN BIGHORN SHEEP IN
THE INTERMOUNTAIN WEST. The Great Basin Naturalist, 51(3),
205–225.
Smith, T. S., Flinders, J. T., & Winn, D. S. (1991b). A HABITAT EVALUATION
PROCEDURE FOR ROCKY MOUNTAIN BIGHORN SHEEP IN
THE INTERMOUNTAIN WEST. The Great Basin Naturalist, 51(3),
205–225.
Soulé, M. E., & Terborgh, J. (1862). Continental Conservation: Scientific
Foundations of Regional Reserve Networks. Island Press.
Stewart, F. E. C., Heim, N. A., Clevenger, A. P., Paczkowski, J., Volpe, J. P.,
& Fisher, J. T. (2016). Wolverine behavior varies spatially with anthropogenic
footprint: implications for conservation and inferences
about declines. Ecology and Evolution, 6(5), 1493–1503.
https://doi.org/10.1002/ece3.1921
Stoner, D. C., Rieth, W. R., Wolfe, M. L., Mecham, M. B., & Neville, A.
(2008). Long-Distance Dispersal of a Female Cougar in a Basin
and Range Landscape. Journal of Wildlife Management, 72(4),
933–939. https://doi.org/10.2193/2007-219
Switalski, T. A., & Nelson, C. R. (2011). Efficacy of road removal for
restoring wildlife habitat: Black bear in the Northern Rocky
Mountains, USA. Biological Conservation, 144(11), 2666–2673.
https://doi.org/10.1016/j.biocon.2011.07.026
Theobald, D. M. (2001). Land-Use Dynamics Beyond the American Urban
Fringe*. Geographical Review, 91(3), 544–564. https://doi.
org/10.1111/j.1931-0846.2001.tb00240.x
Theobald, D. M. (2003). Targeting Conservation Action through Assessment
of Protection and Exurban Threats. Conservation
Biology, 17(6), 1624–1637. https://doi.org/10.1111/j.1523-
1739.2003.00250.x
Theobald, D. M. (2003a). Targeting Conservation Action through Assessment
of Protection and Exurban Threats. Conservation
Biology, 17(6), 1624–1637. https://doi.org/10.1111/j.1523-
1739.2003.00250.x
Theobald, D. M. (2003b). Targeting Conservation Action through Assessment
of Protection and Exurban Threats. Conservation Biology, 17(6), 1624–1637. https://doi.org/10.1111/j.1523-
1739.2003.00250.x
Theobald, D. M. (2004). Placing exurban land-use change in a human
modification framework. Frontiers in Ecology and the
Environment, 2(3), 139–144. https://doi.org/10.1890/1540-
9295(2004)002[0139:PELCIA]2.0.CO;2
Theobald, D. M. (2004a). Placing exurban land-use change in a human
modification framework. Frontiers in Ecology and the
Environment, 2(3), 139–144. https://doi.org/10.1890/1540-
9295(2004)002[0139:PELCIA]2.0.CO;2
Theobald, D. M. (2004b). Placing exurban land‐use change in a human
modification framework. Frontiers in Ecology and the
Environment, 2(3), 139–144. https://doi.org/10.1890/1540-
9295(2004)002[0139:PELCIA]2.0.CO;2
Thompson, D. J., & Jenks, J. A. (2010). Dispersal movements of subadult
cougars from the Black Hills: the notions of range expansion and
recolonization. Ecosphere, 1(4), 1–11. https://doi.org/10.1890/
ES10-00028.1
Thompson, D. J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009a).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, D. J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009b).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, Daniel J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009a).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, Daniel J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009b).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, Daniel J., Jenks, J. A., & Fecske, D. M. (2014). Prevalence
of human-caused mortality in an unhunted cougar population
and potential impacts to management. Wildlife Society Bulletin,
38(2), 341–347. https://doi.org/10.1002/wsb.390
Torres, S. G., Mansfield, T. M., Foley, J. E., Lupo, T., & Brinkhaus, A.
(1996a). Mountain lion and human activity in California: Testing
speculations. Wildlife Society Bulletin, 24(3), 451–460.
Torres, S. G., Mansfield, T. M., Foley, J. E., Lupo, T., & Brinkhaus, A.
(1996b). Mountain lion and human activity in California: Testing
speculations. Wildlife Society Bulletin, 24(3), 451–460.
Travis, W. R. (2013). New Geographies of the American West: Land Use
and the Changing Patterns of Place. Island Press.
Trombulak, S. C., & Frissell, C. A. (2000). Review of Ecological Effects
of Roads on Terrestrial and Aquatic Communities. Conservation
Biology, 14(1), 18–30. https://doi.org/10.1046/j.1523-
1739.2000.99084.x
Turner, B. L., Lambin, E. F., & Reenberg, A. (2007). The emergence of land
change science for global environmental change and sustainability.
Proceedings of the National Academy of Sciences, 104(52),
20666–20671. https://doi.org/10.1073/pnas.0704119104
Vitousek, P. M., Mooney, H. A., Lubchenco, J., & Melillo, J. M. (1997).
Human Domination of Earth’s Ecosystems. Science, 277(5325),
494–499. https://doi.org/10.1126/science.277.5325.494
Wilcove, D. S., Rothstein, D., Dubow, J., Phillips, A., & Losos, E. (1998).
Quantifying Threats to Imperiled Species in the United States.
BioScience, 48(8), 607–615. https://doi.org/10.2307/1313420
Woodroffe, R., & Ginsberg, J. R. (1998). Edge Effects and the Extinction
of Populations Inside Protected Areas. Science, 280(5372),
2126–2128. https://doi.org/10.1126/science.280.5372.2126
Wright, B.S. (1959). The ghost of North America, first edition. Vantange
Press, NewYork, NY, USA.
Wydeven, A. P., Fuller, T. K., Weber, W., & MacDonald, K. (1998). The
Potential for Wolf Recovery in the Northeastern United States
via Dispersal from Southeastern Canada. Wildlife Society Bulletin
(1973-2006), 26(4), 776–784.
Young, S. P., 1889-1969, Goldman, E. A., & 1873-1946. (1946). puma,
mysterious American cat. Retrieved from http://agris.fao.org/
agris-search/search.do?recordID=US201300682670
Young, S. P., 1889-1969, Goldman, E. A., & 1873-1946. (1946). puma,
mysterious American cat. Retrieved from http://agris.fao.org/
agris-search/search.do?recordID=US201300682670
Zarri, A. A., Rahmani, A. R., Singh, A., & Kushwaha, S. P. S. (2008). Habitat
suitability assessment for the endangered Nilgiri Laughingthrush:
A multiple logistic regression approach. Current Science,
94(11), 1487–1494.
Zeigenfuss, L. C., Singer, F. J., & Gudorf, M. A. (2000). Test of a Modified
Habitat Suitability Model for Bighorn Sheep. Restoration
Ecology, 8(4S), 38–46. https://doi.org/10.1046/j.1526-
100x.2000.80064.x
Zhao, C., Li, Y., Li, D., Guan, J., Xiong, Y., & Hu, J. (2014). Habitat suitability
assessment of Sichuan sika deer in Tiebu Nature Reserve during
periods of green and dry grass. Acta Ecologica Sinica, 34(3),
135–140. https://doi.org/10.1016/j.chnaes.2014.03.001
Zhao, C., Li, Y., Li, D., Guan, J., Xiong, Y., & Hu, J. (2014). Habitat suitability
assessment of Sichuan sika deer in Tiebu Nature Reserve during
periods of green and dry grass. Acta Ecologica Sinica, 34(3),
135–140. https://doi.org/10.1016/j.chnaes.2014.03.001
& Kumar, S. (2014). Assessment of suitable habitat of near
threatened striped hyena (Hyaena hyaena Linnaeus, 1758) using
Remote Sensing and Geographic Information System. Asian
Journal of Geoinformatics, 14(2). Retrieved from http://www.
geoinfo.ait.ac.th/ajg/index.php/journal/article/view/139
Alam, M. S., Khan, J. A., Kushwaha, S. P. S., Agrawal, R., Pathak, B. J.,
& Kumar, S. (2014). Assessment of suitable habitat of near
threatened striped hyena (Hyaena hyaena Linnaeus, 1758) using
Remote Sensing and Geographic Information System. Asian
Journal of Geoinformatics, 14(2). Retrieved from http://www.
geoinfo.ait.ac.th/ajg/index.php/journal/article/view/139
Alexander, S. M., Logan, T. B., & Paquet, P. C. (2006). Spatio-temporal
co-occurrence of cougars (Felis concolor), wolves (Canis lupus)
and their prey during winter: a comparison of two analytical
methods. Journal of Biogeography, 33(11), 2001–2012. https://
doi.org/10.1111/j.1365-2699.2006.01564.x
Alexander, S. M., Logan, T. B., & Paquet, P. C. (2006a). Spatio-temporal
co-occurrence of cougars (Felis concolor), wolves (Canis lupus)
and their prey during winter: a comparison of two analytical
methods. Journal of Biogeography, 33(11), 2001–2012. https://
doi.org/10.1111/j.1365-2699.2006.01564.x
Alexander, S. M., Logan, T. B., & Paquet, P. C. (2006b). Spatio-temporal
co-occurrence of cougars (Felis concolor), wolves (Canis lupus)
and their prey during winter: a comparison of two analytical
methods. Journal of Biogeography, 33(11), 2001–2012. https://
doi.org/10.1111/j.1365-2699.2006.01564.x
Alverson, W. S., Waller, D. M., & Solheim, S. L. (1988). Forests Too Deer:
Edge Effects in Northern Wisconsin. Conservation Biology, 2(4),
348–358. https://doi.org/10.1111/j.1523-1739.1988.tb00199.x
Anderson, A.E., (1983). A critical review of literature on Puma (Felis
concolor). Special Report 54. Colorado Division of Wildlife, Fort
Collins, CO, USA.
Apps, C. D., & McLellan, B. N. (2006). Factors influencing the dispersion
and fragmentation of endangered mountain caribou populations.
Biological Conservation, 130(1), 84–97. https://doi.
org/10.1016/j.biocon.2005.12.004
Apps, C. D., & McLellan, B. N. (2006). Factors influencing the dispersion
and fragmentation of endangered mountain caribou populations.
Biological Conservation, 130(1), 84–97. https://doi.
org/10.1016/j.biocon.2005.12.004
Arundel, T., D. Mattson, & Hart. J. (2007). Movements and habitat selection
by mountain lions in the Flagstaff uplands. Pages 17–30
in D. J. Mattson, editor. Mountain lions of the Flagstaff uplands,
2003–2006 progress report. U.S. Geological Survey Washington,
D.C., USA.
Baron, J. S., Theobald, D. M., & Fagre, D. B. (2000). Management of
Land Use Conflicts in the United States Rocky Mountains. Mountain
Research and Development, 20(1), 24–27. https://doi.
org/10.1659/0276-4741(2000)020[0024:MOLUCI]2.0.CO;2
Beier, P. (1993). Determining Minimum Habitat Areas and Habitat Corridors
for Cougars. Conservation Biology, 7(1), 94–108. https://
doi.org/10.1046/j.1523-1739.1993.07010094.x
Beier, P., & Noss, R. F. (1998). Do Habitat Corridors Provide Connectivity?
Conservation Biology, 12(6), 1241–1252. https://doi.
org/10.1111/j.1523-1739.1998.98036.x
Beier, P., Majka, D. R., & Newell, S. L. (2009). Uncertainty analysis of
least-cost modeling for designing wildlife linkages. Ecological
Applications, 19(8), 2067–2077. https://doi.org/10.1890/08-
1898.1
Beier, P., Majka, D. R., & Newell, S. L. (2009). Uncertainty analysis of
least-cost modeling for designing wildlife linkages. Ecological
Applications, 19(8), 2067–2077. https://doi.org/10.1890/08-
1898.1
Bolgiano, C., (1995). Mountain Lion: An Unnatural History of Pumas and
People. Stack-pole Books, Harrisburg, PA, USA.
Burdett, C. L., Crooks, K. R., Theobald, D. M., Wilson, K. R., Boydston,
E. E., Lyren, L. M., … Boyce, W. M. (2010). Interfacing models of
wildlife habitat and human development to predict the future
distribution of puma habitat. Ecosphere, 1(1), 1–21. https://doi.
org/10.1890/ES10-00005.1
Burdett, C. L., K. R. Crooks, D. M. Theobald, K. R. Wilson, E. E.Boydston,
L. M. Lyren, R. N.
Busch, R. H. (2004). The Cougar Almanac: A Complete Natural History of
the Mountain Lion (The Lyons Press: Guildord, CT).
Callan, R., Nibbelink, N. P., Rooney, T. P., Wiedenhoeft, J. E., & Wydeven,
A. P. (2013). Recolonizing wolves trigger a trophic cascade in
Wisconsin (USA). Journal of Ecology, 101(4), 837–845. https://
doi.org/10.1111/1365-2745.12095
Callan, R., Nibbelink, N. P., Rooney, T. P., Wiedenhoeft, J. E., & Wydeven,
A. P. (2013). Recolonizing wolves trigger a trophic cascade in
Wisconsin (USA). Journal of Ecology, 101(4), 837–845. https://
doi.org/10.1111/1365-2745.12095
Červinka, J., Šálek, M., Padyšáková, E., & Šmilauer, P. (2013). The effects
of local and landscape-scale habitat characteristics and prey
availability on corridor use by carnivores: A comparison of two
contrasting farmlands. Journal for Nature Conservation, 21(2),
105–113. https://doi.org/10.1016/j.jnc.2012.11.004
Clevenger, A. P., Wierzchowski, J., Chruszcz, B., & Gunson, K. (2002a).
GIS-Generated, Expert-Based Models for Identifying Wildlife
Habitat Linkages and Planning Mitigation Passages. Conservation
Biology, 16(2), 503–514. https://doi.org/10.1046/j.1523-
1739.2002.00328.x
Clevenger, A. P., Wierzchowski, J., Chruszcz, B., & Gunson, K. (2002b).
GIS-Generated, Expert-Based Models for Identifying Wildlife
Habitat Linkages and Planning Mitigation Passages. Conservation
Biology, 16(2), 503–514. https://doi.org/10.1046/j.1523-
1739.2002.00328.x
Crooks, K.R. and Soulé, M.E. (1999) Mesopredator Release and Avifaunal
Extinctions in a Fragmented System. Nature, 400, 563-566.
http://dx.doi.org/10.1038/23028
Crooks, K. R. (2002). Relative Sensitivities of Mammalian Carnivores to
Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Crooks, K. R. (2002a). Relative Sensitivities of Mammalian Carnivores
to Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Crooks, K. R. (2002b). Relative Sensitivities of Mammalian Carnivores
to Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Crooks, K. R. (2002c). Relative Sensitivities of Mammalian Carnivores
to Habitat Fragmentation. Conservation Biology, 16(2), 488–502.
https://doi.org/10.1046/j.1523-1739.2002.00386.x
Culver, M., Johnson, W. E., Pecon-Slattery, J., & O’Brien, S. J. (2000).
Genomic ancestry of the American puma (Puma concolor).
Journal of Heredity, 91(3), 186–197. https://doi.org/10.1093/
jhered/91.3.186
Cumming, G. S. (2000). Using habitat models to map diversity: pan-
African species richness of ticks (Acari: Ixodida). Journal of
Biogeography, 27(2), 425–440. https://doi.org/10.1046/j.1365-
2699.2000.00419.x
Dickson, B. G., & Beier, P. (2002). Home-Range and Habitat Selection by
Adult Cougars in Southern California. The Journal of Wildlife Management,
66(4), 1235–1245. https://doi.org/10.2307/3802956
Dickson, B. G., Jenness, J. S., Beier, P., & Gehrt. (2005). Influence of vegetation,
topography, and roads on cougar movement in southern california. Journal of Wildlife Management, 69(1), 264–276.
https://doi.org/10.2193/0022-541X(2005)069<0264:IOVTAR>2
.0.CO;2
Drake, N. (2011). Story one: Genes, sightings retrace path of cougar’s
journey: Biologists track cat from the Black Hills to Connecticut.
Science News, 180(5), 5–6. https://doi.org/10.1002/
scin.5591800503
Forman, R. T. T. (1987). The Ethics of Isolation, the Spread of Disturbance,
and Landscape Ecology. In Landscape Heterogeneity and
Disturbance (pp. 213–229). Springer, New York, NY. https://doi.
org/10.1007/978-1-4612-4742-5_12
Fortin, D., Beyer, H. L., Boyce, M. S., Smith, D. W., Duchesne, T., & Mao,
J. S. (2005). Wolves Influence Elk Movements: Behavior Shapes
a Trophic Cascade in Yellowstone National Park. Ecology, 86(5),
1320–1330. https://doi.org/10.1890/04-0953
Gehring, T. M., & Potter, B. A. (2005). Wolf habitat analysis in Michigan:
an example of the need for proactive land management for
carnivore species. Wildlife Society Bulletin, 33(4), 1237–1244.
https://doi.org/10.2193/0091-7648(2005)33[1237:WHAIMA]2.
0.CO;2
Gehring, T. M., & Potter, B. A. (2005). Wolf habitat analysis in Michigan:
an example of the need for proactive land management for
carnivore species. Wildlife Society Bulletin, 33(4), 1237–1244.
https://doi.org/10.2193/0091-7648(2005)33[1237:WHAIMA]2.
0.CO;2
Gilad, O., Wu, X. B., & Armstrong, F. (2013). Assessing the feasibility
for reintroducing desert bighorn sheep to Guadalupe Mountains
National Park: Habitat, migration corridors and challenges.
Applied Geography, 41, 96–104. https://doi.org/10.1016/j.apgeog.
2013.03.012
Gilad, O., Wu, X. B., & Armstrong, F. (2013a). Assessing the feasibility
for reintroducing desert bighorn sheep to Guadalupe Mountains
National Park: Habitat, migration corridors and challenges.
Applied Geography, 41, 96–104. https://doi.org/10.1016/j.apgeog.
2013.03.012
Gilad, O., Wu, X. B., & Armstrong, F. (2013b). Assessing the feasibility
for reintroducing desert bighorn sheep to Guadalupe Mountains
National Park: Habitat, migration corridors and challenges.
Applied Geography, 41, 96–104. https://doi.org/10.1016/j.apgeog.
2013.03.012
Girvetz, E. H., Thorne, J. H., Berry, A. M., & Jaeger, J. A. G. (2008). Integration
of landscape fragmentation analysis into regional planning:
A statewide multi-scale case study from California, USA.
Landscape and Urban Planning, 86(3), 205–218. https://doi.
org/10.1016/j.landurbplan.2008.02.007
Girvetz, E. H., Thorne, J. H., Berry, A. M., & Jaeger, J. A. G. (2008). Integration
of landscape fragmentation analysis into regional planning:
A statewide multi-scale case study from California, USA.
Landscape and Urban Planning, 86(3), 205–218. https://doi.
org/10.1016/j.landurbplan.2008.02.007
Glick, H. B. (2014). Modeling cougar habitat in the Northeastern
United States. Ecological Modelling, 285, 78–89. https://doi.
org/10.1016/j.ecolmodel.2014.02.005
Glick, H. B. (2014). Modeling cougar habitat in the Northeastern
United States. Ecological Modelling, 285, 78–89. https://doi.
org/10.1016/j.ecolmodel.2014.02.005
Gompper, M. E. (2002). Top Carnivores in the Suburbs? Ecological and
Conservation Issues Raised by Colonization of North-eastern
North America by CoyotesThe expansion of the coyote’s geographical
range may broadly influence community structure, and
rising coyote densities in the suburbs may alter how the general
public views wildlife. BioScience, 52(2), 185–190. https://doi.
org/10.1641/0006-3568(2002)052[0185:TCITSE]2.0.CO;2
Gray, M., Wilmers, C. C., Reed, S. E., & Merenlender, A. M. (2016a).
Landscape feature-based permeability models relate to puma
occurrence. Landscape and Urban Planning, 147, 50–58. https://
doi.org/10.1016/j.landurbplan.2015.11.009
Gray, M., Wilmers, C. C., Reed, S. E., & Merenlender, A. M. (2016b).
Landscape feature-based permeability models relate to puma
occurrence. Landscape and Urban Planning, 147, 50–58. https://
doi.org/10.1016/j.landurbplan.2015.11.009
Hansen, A. J., Rasker, R., Maxwell, B., Rotella, J. J., Johnson, J. D.,
Parmenter, A. W., … Kraska, M.P. V. (2002). Ecological Causes
and Consequences of Demographic Change in the New West.
BioScience, 52(2), 151–162. https://doi.org/10.1641/0006-
3568(2002)052[0151:ECACOD]2.0.CO;2
Harveson, L.A., Harveson, P.M. & Adams, R.W.(2003). Proceedings of
the Sixth Mountain Lion Workshop , Austin, Texas, USA.
Hénaux, V., Powell, L. A., Hobson, K. A., Nielsen, C. K., & LaRue, M. A.
(2011). Tracking large carnivore dispersal using isotopic clues
in claws: an application to cougars across the Great Plains.
Methods in Ecology and Evolution, 2(5), 489–499. https://doi.
org/10.1111/j.2041-210X.2011.00107.x
Hénaux, V., Powell, L. A., Hobson, K. A., Nielsen, C. K., & LaRue, M. A.
(2011). Tracking large carnivore dispersal using isotopic clues
in claws: an application to cougars across the Great Plains.
Methods in Ecology and Evolution, 2(5), 489–499. https://doi.
org/10.1111/j.2041-210X.2011.00107.x
Holmes, B. R., & Laundré, J. W. (2006). Use of open, edge and forest
areas by pumas Puma concolor in winter: are pumas foraging
optimally? Wildlife Biology, 12(2), 201–209. https://doi.
org/10.2981/0909-6396(2006)12[201:UOOEAF]2.0.CO;2
Hortal, J., Lobo, J. M., & Martín-piera, F. (2001). Forecasting insect
species richness scores in poorly surveyed territories: the
case of the Portuguese dung beetles (Col. Scarabaeinae).
Biodiversity & Conservation, 10(8), 1343–1367. https://doi.
org/10.1023/A:1016624500023
Hunter, R. D., Fisher, R. N., & Crooks, K. R. (2003). Landscape-level
connectivity in coastal southern California, USA, as assessed
through carnivore habitat suitability. Natural Areas Journal,
23(4), 302314.
Jenkins, M. (2003a). Prospects for Biodiversity. Science, 302(5648),
1175–1177. https://doi.org/10.1126/science.1088666
Jenkins, M. (2003b). Prospects for Biodiversity. Science, 302(5648),
1175–1177. https://doi.org/10.1126/science.1088666
Keeley, A. T. H., Beier, P., & Gagnon, J. W. (2016). Estimating landscape
resistance from habitat suitability: effects of data source and nonlinearities. Landscape Ecology, 31(9), 2151–2162. https://
doi.org/10.1007/s10980-016-0387-5
Kertson, B. N., Spencer, R. D., Marzluff, J. M., Hepinstall-Cymerman, J.,
& Grue, C. E. (2011). Cougar space use and movements in the
wildland–urban landscape of western Washington. Ecological
Applications, 21(8), 2866–2881. https://doi.org/10.1890/11-
0947.1
Knopff, A. A., Knopff, K. H., Boyce, M. S., & St. Clair, C. C. (2014). Flexible
habitat selection by cougars in response to anthropogenic development.
Biological Conservation, 178, 136–145. https://doi.
org/10.1016/j.biocon.2014.07.017
Knopff, A. A., Knopff, K. H., Boyce, M. S., & St. Clair, C. C. (2014). Flexible
habitat selection by cougars in response to anthropogenic development.
Biological Conservation, 178, 136–145. https://doi.
org/10.1016/j.biocon.2014.07.017
Kushwaha, S. P. S., Khan, A., Habib, B., Quadri, A., & Singh, A. (2004).
Evaluation of sambar and muntjak habitats using geostatistical
modelling. Current Science, 86(10), 1390–1400.
LaRue, M. A., & Nielsen, C. K. (2008). Modelling potential dispersal corridors
for cougars in midwestern North America using least-cost
path methods. Ecological Modelling, 212(3), 372–381. https://
doi.org/10.1016/j.ecolmodel.2007.10.036
LaRue, M. A., & Nielsen, C. K. (2008). Modelling potential dispersal corridors
for cougars in midwestern North America using least-cost
path methods. Ecological Modelling, 212(3), 372–381. https://
doi.org/10.1016/j.ecolmodel.2007.10.036
LaRue, M. A., & Nielsen, C. K. (2016). Population viability of recolonizing
cougars in midwestern North America. Ecological Modelling, 321,
121–129. https://doi.org/10.1016/j.ecolmodel.2015.09.026
LaRue, M. A., & Nielsen, C. K. (2016a). Population viability of recolonizing
cougars in midwestern North America. Ecological
Modelling, 321, 121–129. https://doi.org/10.1016/j.ecolmodel.
2015.09.026
LaRue, M. A., & Nielsen, C. K. (2016b). Population viability of recolonizing
cougars in midwestern North America. Ecological
Modelling, 321, 121–129. https://doi.org/10.1016/j.ecolmodel.
2015.09.026
LaRue, M. A., Nielsen, C. K., Dowling, M., Miller, K., Wilson, B., Shaw,
H., & Anderson, C. R. (2012). Cougars are recolonizing the midwest:
Analysis of cougar confirmations during 1990–2008. The
Journal of Wildlife Management, 76(7), 1364–1369. https://doi.
org/10.1002/jwmg.396
Laundré, J. W., & Loxterman, J. (2007). Impact of Edge Habitat on Summer
Home Range Size in Female Pumas. The American Midland
Naturalist, 157(1), 221–229. https://doi.org/10.1674/0003-
0031(2007)157[221:IOEHOS]2.0.CO;2
Lenton, S. M., Fa, J. E., & Val, J. P. D. (2000). A simple non-parametric GIS
model for predicting species distribution: endemic birds in Bioko
Island, West Africa. Biodiversity & Conservation, 9(7), 869–885.
https://doi.org/10.1023/A:1008980910283
Light, J. T. (1971). An ecological view of bighorn habitat on Mt. San Antonio.
Transactions of North American wild sheep Conference 1,
pp. 150-157.
Liu, S., Dong, Y., Cheng, F., Zhang, Y., Hou, X., Dong, S., & Coxixo, A.
(2017). Effects of road network on Asian elephant habitat and
connectivity between the nature reserves in Xishuangbanna,
Southwest China. Journal for Nature Conservation, 38, 11–20.
https://doi.org/10.1016/j.jnc.2017.05.001
Liu, S., Dong, Y., Cheng, F., Zhang, Y., Hou, X., Dong, S., & Coxixo, A.
(2017). Effects of road network on Asian elephant habitat and
connectivity between the nature reserves in Xishuangbanna,
Southwest China. Journal for Nature Conservation, 38, 11–20.
https://doi.org/10.1016/j.jnc.2017.05.001
Logan, K. A., & Irwin, L. L. (1985). Mountain Lion Habitats in the Big
Horn Mountains, Wyoming. Wildlife Society Bulletin (1973-
2006), 13(3), 257–262.
Logan, K. A., & Sweanor, L. L. (2001). Desert Puma: Evolutionary Ecology
And Conservation Of An Enduring Carnivore. Island Press.
Maehr, D. S., Land, E. D., Shindle, D. B., Bass, O. L., & Hoctor, T. S.
(2002). Florida panther dispersal and conservation. Biological
Conservation, 106(2), 187–197. https://doi.org/10.1016/S0006-
3207(01)00245-2
Marzluff, J. M., & Ewing, K. (2001). Restoration of Fragmented Landscapes
for the Conservation of Birds: A General Framework and
Specific Recommendations for Urbanizing Landscapes. Restoration
Ecology, 9(3), 280–292. https://doi.org/10.1046/j.1526-
100x.2001.009003280.x
Marzluff, J. M., & Ewing, K. (2008). Restoration of Fragmented Landscapes
for the Conservation of Birds: A General Framework and
Specific Recommendations for Urbanizing Landscapes. In Urban
Ecology (pp. 739–755). Springer, Boston, MA. https://doi.
org/10.1007/978-0-387-73412-5_48
Morrison, C. D., Boyce, M. S., Nielsen, S. E., & Bacon, M. M. (2014). Habitat
selection of a re-colonized cougar population in response to
seasonal fluctuations of human activity. The Journal of Wildlife
Management, 78(8), 1394–1403. https://doi.org/10.1002/
jwmg.799
Morrison, C. D., Boyce, M. S., Nielsen, S. E., & Bacon, M. M. (2014). Habitat
selection of a re-colonized cougar population in response to
seasonal fluctuations of human activity. The Journal of Wildlife
Management, 78(8), 1394–1403. https://doi.org/10.1002/
jwmg.799
Morrison, S. A., & Boyce, W. M. (2009). Conserving Connectivity: Some
Lessons from Mountain Lions in Southern California. Conservation
Biology, 23(2), 275–285. https://doi.org/10.1111/j.1523-
1739.2008.01079.x
Northrup, J. M., Pitt, J., Muhly, T. B., Stenhouse, G. B., Musiani, M., &
Boyce, M. S. (2012). Vehicle traffic shapes grizzly bear behaviour
on a multiple-use landscape. Journal of Applied Ecology, 49(5),
1159–1167. https://doi.org/10.1111/j.1365-2664.2012.02180.x
Noss, R. F., & Daly K. M. (2006). Incorporating connectivity into broad
scale conservation planning. Pages 587–619 in K. R. Crooks and
M. A. Sanjayan, editors. Connectivity conservation, Cambridge
University Press, Cambridge, United Kingdom
Noss, R. F., Quigley, H. B., Hornocker, M. G., Merrill, T., & Paquet, P.
C. (1996). Conservation Biology and Carnivore Conservation in
the Rocky Mountains. Conservation Biology, 10(4), 949–963.
https://doi.org/10.1046/j.1523-1739.1996.10040949.x
O′Neil, S. T., Rahn, K. C., & Bump, J. K. (2014). Habitat Capacity for Cougar
Recolonization in the Upper Great Lakes Region. PLOS ONE,
9(11), e112565. https://doi.org/10.1371/journal.pone.0112565
Pletscher, D. H., Ream, R. R., Boyd, D. K., Fairchild, M. W., & Kunkel, K. E.
(1997). Population Dynamics of a Recolonizing Wolf Population.
The Journal of Wildlife Management, 61(2), 459–465. https://
doi.org/10.2307/3802604
Pyare, S., Cain, S., Moody, D., Schwartz, C., & Berger, J. (2004). Carnivore
re-colonisation: reality, possibility and a non-equilibrium century
for grizzly bears in the southern Yellowstone ecosystem. Animal
Conservation Forum, 7(1), 71–77. https://doi.org/10.1017/
S1367943003001203
Restas, A. (2015). Drone Applications for Supporting Disaster Management.
World Journal of Engineering and Technology, 03(03),
316–321. https://doi.org/10.4236/wjet.2015.33C047
Ripple, W. J., & Beschta, R. L. (2004). Wolves and the Ecology of Fear: Can
Predation Risk Structure Ecosystems? BioScience, 54(8), 755–
766. https://doi.org/10.1641/0006-3568(2004)054[0755:WATE
OF]2.0.CO;2
Ripple, W. J., & Beschta, R. L. (2004a). Wolves and the Ecology of Fear:
Can Predation Risk Structure Ecosystems? BioScience, 54(8), 755–
766. https://doi.org/10.1641/0006-3568(2004)054[0755:WATE
OF]2.0.CO;2
Ripple, W. J., & Beschta, R. L. (2004b). Wolves and the Ecology of Fear:
Can Predation Risk Structure Ecosystems? BioScience, 54(8), 755–
766. https://doi.org/10.1641/0006-3568(2004)054[0755:WATE
OF]2.0.CO;2
Ripple, W. J., & Beschta, R. L. (2006a). Linking a cougar decline, trophic
cascade, and catastrophic regime shift in Zion National
Park. Biological Conservation, 133(4), 397–408. https://doi.
org/10.1016/j.biocon.2006.07.002
Ripple, W. J., & Beschta, R. L. (2006b). Linking a cougar decline, trophic
cascade, and catastrophic regime shift in Zion National
Park. Biological Conservation, 133(4), 397–408. https://doi.
org/10.1016/j.biocon.2006.07.002
Ripple, W. J., & Beschta, R. L. (2008). Trophic cascades involving cougar,
mule deer, and black oaks in Yosemite National Park. Biological
Conservation, 141(5), 1249–1256. https://doi.org/10.1016/j.
biocon.2008.02.028
Rosatte, R. (2011). Evidence Confirms the Presence of Cougars ( Puma
concolor ) in Ontario, Canada. The Canadian Field-Naturalist,
125(2), 116–125.
Saaty, T.L. (1980). The Analytical Hierarchy Process: Planning, Setting
Priorities, Resource Allocation. McGraw-Hill International Book
Co., New York.
Sanderson, E. W., Redford, K. H., Vedder, A., Coppolillo, P. B., & Ward,
S. E. (2002a). A conceptual model for conservation planning
based on landscape species requirements. Landscape and Urban
Planning, 58(1), 41–56. https://doi.org/10.1016/S0169-
2046(01)00231-6
Sanderson, E. W., Redford, K. H., Vedder, A., Coppolillo, P. B., & Ward,
S. E. (2002b). A conceptual model for conservation planning
based on landscape species requirements. Landscape and Urban
Planning, 58(1), 41–56. https://doi.org/10.1016/S0169-
2046(01)00231-6
Schamberger, M., & Krohn, W. (1982). Status of the Habitat Evaluation
Procedures. US Fish & Wildlife Publications. Retrieved from
https://digitalcommons.unl.edu/usfwspubs/48
Schipper, J., Chanson, J. S., Chiozza, F., Cox, N. A., Hoffmann, M., Katariya,
V., … Young, B. E. (2008). The Status of the World’s Land and
Marine Mammals: Diversity, Threat, and Knowledge. Science,
322(5899), 225–230. https://doi.org/10.1126/science.1165115
Shen, G., Feng, C., Xie, Z., Ouyang, Z., Li, J., & Pascal, M. (2008). Proposed
Conservation Landscape for Giant Pandas in the Minshan
Mountains, China. Conservation Biology, 22(5), 1144–1153.
https://doi.org/10.1111/j.1523-1739.2008.01038.x
Singer, F. J., Bleich, V. C., & Gudorf, M. A. (2000). Restoration of Bighorn
Sheep Metapopulations in and Near Western National Parks.
Restoration Ecology, 8(4S), 14–24. https://doi.org/10.1046/
j.1526-100x.2000.80062.x
Smith, T. S., & Flinders, J. T. (1992). Evaluation of mountain sheep habitat
in Zion National Park, Utah. Desert Bighorn Council Transactions,
36, 4-9.
Smith, T. S., Flinders, J. T., & Winn, D. S. (1991a). A HABITAT EVALUATION
PROCEDURE FOR ROCKY MOUNTAIN BIGHORN SHEEP IN
THE INTERMOUNTAIN WEST. The Great Basin Naturalist, 51(3),
205–225.
Smith, T. S., Flinders, J. T., & Winn, D. S. (1991b). A HABITAT EVALUATION
PROCEDURE FOR ROCKY MOUNTAIN BIGHORN SHEEP IN
THE INTERMOUNTAIN WEST. The Great Basin Naturalist, 51(3),
205–225.
Soulé, M. E., & Terborgh, J. (1862). Continental Conservation: Scientific
Foundations of Regional Reserve Networks. Island Press.
Stewart, F. E. C., Heim, N. A., Clevenger, A. P., Paczkowski, J., Volpe, J. P.,
& Fisher, J. T. (2016). Wolverine behavior varies spatially with anthropogenic
footprint: implications for conservation and inferences
about declines. Ecology and Evolution, 6(5), 1493–1503.
https://doi.org/10.1002/ece3.1921
Stoner, D. C., Rieth, W. R., Wolfe, M. L., Mecham, M. B., & Neville, A.
(2008). Long-Distance Dispersal of a Female Cougar in a Basin
and Range Landscape. Journal of Wildlife Management, 72(4),
933–939. https://doi.org/10.2193/2007-219
Switalski, T. A., & Nelson, C. R. (2011). Efficacy of road removal for
restoring wildlife habitat: Black bear in the Northern Rocky
Mountains, USA. Biological Conservation, 144(11), 2666–2673.
https://doi.org/10.1016/j.biocon.2011.07.026
Theobald, D. M. (2001). Land-Use Dynamics Beyond the American Urban
Fringe*. Geographical Review, 91(3), 544–564. https://doi.
org/10.1111/j.1931-0846.2001.tb00240.x
Theobald, D. M. (2003). Targeting Conservation Action through Assessment
of Protection and Exurban Threats. Conservation
Biology, 17(6), 1624–1637. https://doi.org/10.1111/j.1523-
1739.2003.00250.x
Theobald, D. M. (2003a). Targeting Conservation Action through Assessment
of Protection and Exurban Threats. Conservation
Biology, 17(6), 1624–1637. https://doi.org/10.1111/j.1523-
1739.2003.00250.x
Theobald, D. M. (2003b). Targeting Conservation Action through Assessment
of Protection and Exurban Threats. Conservation Biology, 17(6), 1624–1637. https://doi.org/10.1111/j.1523-
1739.2003.00250.x
Theobald, D. M. (2004). Placing exurban land-use change in a human
modification framework. Frontiers in Ecology and the
Environment, 2(3), 139–144. https://doi.org/10.1890/1540-
9295(2004)002[0139:PELCIA]2.0.CO;2
Theobald, D. M. (2004a). Placing exurban land-use change in a human
modification framework. Frontiers in Ecology and the
Environment, 2(3), 139–144. https://doi.org/10.1890/1540-
9295(2004)002[0139:PELCIA]2.0.CO;2
Theobald, D. M. (2004b). Placing exurban land‐use change in a human
modification framework. Frontiers in Ecology and the
Environment, 2(3), 139–144. https://doi.org/10.1890/1540-
9295(2004)002[0139:PELCIA]2.0.CO;2
Thompson, D. J., & Jenks, J. A. (2010). Dispersal movements of subadult
cougars from the Black Hills: the notions of range expansion and
recolonization. Ecosphere, 1(4), 1–11. https://doi.org/10.1890/
ES10-00028.1
Thompson, D. J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009a).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, D. J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009b).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, Daniel J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009a).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, Daniel J., Fecske, D. M., Jenks, J. A., & Jarding, A. R. (2009b).
Food Habits of Recolonizing Cougars in the Dakotas: Prey Obtained
from Prairie and Agricultural Habitats. The American Midland
Naturalist, 161(1), 69–75. https://doi.org/10.1674/0003-
0031-161.1.69
Thompson, Daniel J., Jenks, J. A., & Fecske, D. M. (2014). Prevalence
of human-caused mortality in an unhunted cougar population
and potential impacts to management. Wildlife Society Bulletin,
38(2), 341–347. https://doi.org/10.1002/wsb.390
Torres, S. G., Mansfield, T. M., Foley, J. E., Lupo, T., & Brinkhaus, A.
(1996a). Mountain lion and human activity in California: Testing
speculations. Wildlife Society Bulletin, 24(3), 451–460.
Torres, S. G., Mansfield, T. M., Foley, J. E., Lupo, T., & Brinkhaus, A.
(1996b). Mountain lion and human activity in California: Testing
speculations. Wildlife Society Bulletin, 24(3), 451–460.
Travis, W. R. (2013). New Geographies of the American West: Land Use
and the Changing Patterns of Place. Island Press.
Trombulak, S. C., & Frissell, C. A. (2000). Review of Ecological Effects
of Roads on Terrestrial and Aquatic Communities. Conservation
Biology, 14(1), 18–30. https://doi.org/10.1046/j.1523-
1739.2000.99084.x
Turner, B. L., Lambin, E. F., & Reenberg, A. (2007). The emergence of land
change science for global environmental change and sustainability.
Proceedings of the National Academy of Sciences, 104(52),
20666–20671. https://doi.org/10.1073/pnas.0704119104
Vitousek, P. M., Mooney, H. A., Lubchenco, J., & Melillo, J. M. (1997).
Human Domination of Earth’s Ecosystems. Science, 277(5325),
494–499. https://doi.org/10.1126/science.277.5325.494
Wilcove, D. S., Rothstein, D., Dubow, J., Phillips, A., & Losos, E. (1998).
Quantifying Threats to Imperiled Species in the United States.
BioScience, 48(8), 607–615. https://doi.org/10.2307/1313420
Woodroffe, R., & Ginsberg, J. R. (1998). Edge Effects and the Extinction
of Populations Inside Protected Areas. Science, 280(5372),
2126–2128. https://doi.org/10.1126/science.280.5372.2126
Wright, B.S. (1959). The ghost of North America, first edition. Vantange
Press, NewYork, NY, USA.
Wydeven, A. P., Fuller, T. K., Weber, W., & MacDonald, K. (1998). The
Potential for Wolf Recovery in the Northeastern United States
via Dispersal from Southeastern Canada. Wildlife Society Bulletin
(1973-2006), 26(4), 776–784.
Young, S. P., 1889-1969, Goldman, E. A., & 1873-1946. (1946). puma,
mysterious American cat. Retrieved from http://agris.fao.org/
agris-search/search.do?recordID=US201300682670
Young, S. P., 1889-1969, Goldman, E. A., & 1873-1946. (1946). puma,
mysterious American cat. Retrieved from http://agris.fao.org/
agris-search/search.do?recordID=US201300682670
Zarri, A. A., Rahmani, A. R., Singh, A., & Kushwaha, S. P. S. (2008). Habitat
suitability assessment for the endangered Nilgiri Laughingthrush:
A multiple logistic regression approach. Current Science,
94(11), 1487–1494.
Zeigenfuss, L. C., Singer, F. J., & Gudorf, M. A. (2000). Test of a Modified
Habitat Suitability Model for Bighorn Sheep. Restoration
Ecology, 8(4S), 38–46. https://doi.org/10.1046/j.1526-
100x.2000.80064.x
Zhao, C., Li, Y., Li, D., Guan, J., Xiong, Y., & Hu, J. (2014). Habitat suitability
assessment of Sichuan sika deer in Tiebu Nature Reserve during
periods of green and dry grass. Acta Ecologica Sinica, 34(3),
135–140. https://doi.org/10.1016/j.chnaes.2014.03.001
Zhao, C., Li, Y., Li, D., Guan, J., Xiong, Y., & Hu, J. (2014). Habitat suitability
assessment of Sichuan sika deer in Tiebu Nature Reserve during
periods of green and dry grass. Acta Ecologica Sinica, 34(3),
135–140. https://doi.org/10.1016/j.chnaes.2014.03.001
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How to Cite
Mbuh, M. J., & Vruno, N. (2018). Habitat Suitability Analysis for Mountain Lions (Puma Concolor) Recolonization/ Reintroduction in Minnesota. European Journal of Ecology, 4(2), 28-40. https://doi.org/10.2478/eje-2018-0013
